Standout Papers

WGCNA: an R package for weighted correlation network analysis 2005 2026 2012 2019 15.9k
  1. WGCNA: an R package for weighted correlation network analysis (2008)
    Peter Langfelder, Steve Horvath BMC Bioinformatics
  2. DNA methylation age of human tissues and cell types (2013)
    Steve Horvath Genome biology
  3. A General Framework for Weighted Gene Co-Expression Network Analysis (2005)
    Bin Zhang, Steve Horvath Statistical Applications in Genetics and Molecular Biology
  4. An epigenetic biomarker of aging for lifespan and healthspan (2018)
    Morgan E. Levine, Ake T. Lu et al. Aging
  5. DNA methylation-based biomarkers and the epigenetic clock theory of ageing (2018)
    Steve Horvath, Kenneth Raj Nature Reviews Genetics
  6. Transcriptomic analysis of autistic brain reveals convergent molecular pathology (2011)
    Irina Voineagu, Xinchen Wang et al. Nature
  7. Defining clusters from a hierarchical cluster tree: the Dynamic Tree Cut package for R (2007)
    Peter Langfelder, Bin Zhang et al. Bioinformatics
  8. DNA methylation GrimAge strongly predicts lifespan and healthspan (2019)
    Ake T. Lu, Austin Quach et al. Aging
  9. Global histone modification patterns predict risk of prostate cancer recurrence (2005)
    David B. Seligson, Steve Horvath et al. Nature
  10. FastRFunctions for Robust Correlations and Hierarchical Clustering (2012)
    Peter Langfelder, Steve Horvath Journal of Statistical Software
  11. Genetic programs in human and mouse early embryos revealed by single-cell RNA sequencing (2013)
    Zhigang Xue, Kevin Huang et al. Nature
  12. Implementing a unified approach to family-based tests of association (2000)
    Nan M. Laird, Steve Horvath et al. Genetic Epidemiology
  13. Integrative Functional Genomic Analyses Implicate Specific Molecular Pathways and Circuits in Autism (2013)
    Neelroop Parikshak, Rui Luo et al. Cell
  14. Fast R Functions for Robust Correlations and Hierarchical Clustering. (2012)
    Peter Langfelder, Steve Horvath PubMed
  15. Epigenetic Predictor of Age (2011)
    Sven Bocklandt, Wen Lin et al. PLoS ONE
  16. Is My Network Module Preserved and Reproducible? (2011)
    Peter Langfelder, Rui Luo et al. PLoS Computational Biology
  17. Geometric Interpretation of Gene Coexpression Network Analysis (2008)
    Steve Horvath, Jun Dong PLoS Computational Biology
  18. The family based association test method: strategies for studying general genotype–phenotype associations (2001)
    Steve Horvath, Xin Xu et al. European Journal of Human Genetics
  19. Eigengene networks for studying the relationships between co-expression modules (2007)
    Peter Langfelder, Steve Horvath BMC Systems Biology
  20. Gene Expression Profiling of Gliomas Strongly Predicts Survival (2004)
    William A. Freije, F. Edmundo Castro-Vargas et al. Cancer Research
  21. Functional organization of the transcriptome in human brain (2008)
    Michael C. Oldham, Geneviève Konopka et al. Nature Neuroscience
  22. Carbonic anhydrase IX is an independent predictor of survival in advanced renal clear cell carcinoma: implications for prognosis and therapy. (2003)
    Matthew Bui, David B. Seligson et al. PubMed
  23. Obesity accelerates epigenetic aging of human liver (2014)
    Steve Horvath, Wiebke Erhart et al. Proceedings of the National Academy of Sciences
  24. Aging effects on DNA methylation modules in human brain and blood tissue (2012)
    Steve Horvath, Yafeng Zhang et al. Genome biology
  25. Genome-wide changes in lncRNA, splicing, and regional gene expression patterns in autism (2016)
    Neelroop Parikshak, Vivek Swarup et al. Nature
  26. The epigenetic clock is correlated with physical and cognitive fitness in the Lothian Birth Cohort 1936 (2015)
    Riccardo E. Marioni, Sonia Shah et al. International Journal of Epidemiology
  27. HIV-1 Infection Accelerates Age According to the Epigenetic Clock (2015)
    Steve Horvath, Andrew J. Levine The Journal of Infectious Diseases
  28. A new aging measure captures morbidity and mortality risk across diverse subpopulations from NHANES IV: A cohort study (2018)
    Zuyun Liu, Pei‐Lun Kuo et al. PLoS Medicine
  29. Epigenetic age of the pre-frontal cortex is associated with neuritic plaques, amyloid load, and Alzheimer’s disease related cognitive functioning (2015)
    Morgan E. Levine, Ake T Lu et al. Aging
  30. Reversal of epigenetic aging and immunosenescent trends in humans (2019)
    Gregory M. Fahy, Robert T. Brooke et al. Aging Cell
  31. GrimAge Outperforms Other Epigenetic Clocks in the Prediction of Age-Related Clinical Phenotypes and All-Cause Mortality (2020)
    Cathal McCrory, Giovanni Fiorito et al. The Journals of Gerontology Series A
  32. Long-term maturation of human cortical organoids matches key early postnatal transitions (2021)
    Aaron Gordon, Se‐Jin Yoon et al. Nature Neuroscience
  33. Making sense of the ageing methylome (2022)
    Kirsten Seale, Steve Horvath et al. Nature Reviews Genetics
  34. Biological age is increased by stress and restored upon recovery (2023)
    Jesse R. Poganik, Bohan Zhang et al. Cell Metabolism
  35. Causality-enriched epigenetic age uncouples damage and adaptation (2024)
    Kejun Ying, Andrei E. Tarkhov et al. Nature Aging
  36. Epigenetic ageing clocks: statistical methods and emerging computational challenges (2025)
    Andrew E. Teschendorff, Steve Horvath Nature Reviews Genetics
  37. Individual and additive effects of vitamin D, omega-3 and exercise on DNA methylation clocks of biological aging in older adults from the DO-HEALTH trial (2025)
    Heike A. Bischoff‐Ferrari, Maud Wieczorek et al. Nature Aging

Immediate Impact

3 by Nobel laureates 46 from Science/Nature 75 standout
Sub-graph 1 of 23

Citing Papers

Antisense oligonucleotide therapeutic approach for Timothy syndrome
2024 StandoutNature
Single-cell brain organoid screening identifies developmental defects in autism
2023 StandoutNature
12 intermediate papers

Works of Steve Horvath being referenced

Integrative Functional Genomic Analyses Implicate Specific Molecular Pathways and Circuits in Autism
2013 Standout
Transcriptomic analysis of autistic brain reveals convergent molecular pathology
2011 StandoutNature
and 6 more

Author Peers

Author Last Decade Papers Cites
Steve Horvath 42656 10762 5130 8104 437 67.8k
Andrew P. Feinberg 44543 15391 5571 6693 233 61.5k
Yoav Benjamini 29933 12221 2880 5338 139 97.8k
Rafael A. Irizarry 37896 7974 1997 7932 181 55.5k
Francis S. Collins 30338 12917 1729 4553 412 59.7k
Trey Ideker 44414 6165 1081 7931 236 63.2k
Manel Esteller 56762 9131 2248 20419 651 73.1k
Li Zhang 27883 4274 6409 7237 1.8k 58.3k
Jin Li 28473 14384 1962 6885 2.5k 70.3k
Peter A. Jones 48140 8776 2466 10709 350 60.4k
Aldons J. Lusis 30239 8896 1198 5104 615 62.5k

All Works

Loading papers...

Rankless by CCL
2026