Penn Lloyd

2.1k total citations
56 papers, 1.6k citations indexed

About

Penn Lloyd is a scholar working on Ecology, Ecology, Evolution, Behavior and Systematics and Nature and Landscape Conservation. According to data from OpenAlex, Penn Lloyd has authored 56 papers receiving a total of 1.6k indexed citations (citations by other indexed papers that have themselves been cited), including 45 papers in Ecology, 28 papers in Ecology, Evolution, Behavior and Systematics and 21 papers in Nature and Landscape Conservation. Recurrent topics in Penn Lloyd's work include Avian ecology and behavior (37 papers), Animal Behavior and Reproduction (23 papers) and Wildlife Ecology and Conservation (20 papers). Penn Lloyd is often cited by papers focused on Avian ecology and behavior (37 papers), Animal Behavior and Reproduction (23 papers) and Wildlife Ecology and Conservation (20 papers). Penn Lloyd collaborates with scholars based in South Africa, United States and United Kingdom. Penn Lloyd's co-authors include Thomas E. Martin, Alina M. Niklison, Ronald D. Bassar, Sonya K. Auer, Martin H. Villet, Rauri C. K. Bowie, Riccardo Ton, Ângela M. Ribeiro, Timothy M. Crowe and R.M. Little and has published in prestigious journals such as PLoS ONE, The American Naturalist and Evolution.

In The Last Decade

Penn Lloyd

54 papers receiving 1.5k citations

Peers — A (Enhanced Table)

Peers by citation overlap · career bar shows stage (early→late) cites · hero ref

Name h Career Trend Papers Cites
Penn Lloyd South Africa 21 1.3k 852 399 212 211 56 1.6k
Pär Forslund Sweden 22 1.5k 1.2× 1.2k 1.4× 345 0.9× 139 0.7× 274 1.3× 36 1.9k
Yolanda E. Morbey Canada 19 930 0.7× 638 0.7× 637 1.6× 130 0.6× 209 1.0× 65 1.5k
Vicente García‐Navas Spain 23 824 0.6× 796 0.9× 268 0.7× 183 0.9× 234 1.1× 72 1.3k
Kim Jaatinen Finland 22 1.0k 0.8× 615 0.7× 239 0.6× 255 1.2× 152 0.7× 61 1.4k
Volker Salewski Germany 24 1.3k 1.0× 655 0.8× 411 1.0× 488 2.3× 171 0.8× 63 1.6k
Kari Koivula Finland 25 1.3k 1.0× 1.0k 1.2× 283 0.7× 162 0.8× 203 1.0× 65 1.6k
Chris M. Perrins United Kingdom 14 1.1k 0.9× 757 0.9× 310 0.8× 468 2.2× 227 1.1× 18 1.6k
Ronald D. Bassar United States 23 1.3k 1.0× 987 1.2× 978 2.5× 204 1.0× 497 2.4× 54 2.1k
Francisco Pulido Germany 18 1.3k 1.0× 800 0.9× 374 0.9× 563 2.7× 391 1.9× 31 1.7k
Patrik Karell Finland 20 916 0.7× 753 0.9× 210 0.5× 208 1.0× 122 0.6× 53 1.3k

Countries citing papers authored by Penn Lloyd

Since Specialization
Citations

This map shows the geographic impact of Penn Lloyd's research. It shows the number of citations coming from papers published by authors working in each country. You can also color the map by specialization and compare the number of citations received by Penn Lloyd with the expected number of citations based on a country's size and research output (numbers larger than one mean the country cites Penn Lloyd more than expected).

Fields of papers citing papers by Penn Lloyd

Since Specialization
Physical SciencesHealth SciencesLife SciencesSocial Sciences

This network shows the impact of papers produced by Penn Lloyd. Nodes represent research fields, and links connect fields that are likely to share authors. Colored nodes show fields that tend to cite the papers produced by Penn Lloyd. The network helps show where Penn Lloyd may publish in the future.

Co-authorship network of co-authors of Penn Lloyd

This figure shows the co-authorship network connecting the top 25 collaborators of Penn Lloyd. A scholar is included among the top collaborators of Penn Lloyd based on the total number of citations received by their joint publications. Widths of edges represent the number of papers authors have co-authored together. Node borders signify the number of papers an author published with Penn Lloyd. Penn Lloyd is excluded from the visualization to improve readability, since they are connected to all nodes in the network.

All Works

20 of 20 papers shown
1.
Lloyd, Penn, et al.. (2025). Habitat associations of the Southern Squatter Pigeon Geophaps scripta scripta in Queensland. Australian field ornithology. 42. 156–167.
2.
Boyce, Andy J., et al.. (2020). Metabolic rate is negatively linked to adult survival but does not explain latitudinal differences in songbirds. Ecology Letters. 23(4). 642–652. 22 indexed citations
3.
Lloyd, Penn, et al.. (2019). Survival synchronicity in two avian insectivore communities. Ibis. 162(3). 787–800. 2 indexed citations
4.
Bosque, Carlos & Penn Lloyd. (2017). Diet and time-activity budget of White-backed MousebirdsColius coliusin south-western South Africa. Ostrich. 88(3). 247–252. 2 indexed citations
5.
Martin, Thomas E., et al.. (2015). Postnatal Growth Rates Covary Weakly with Embryonic Development Rates and Do Not Explain Adult Mortality Probability among Songbirds on Four Continents. The American Naturalist. 185(3). 380–389. 35 indexed citations
6.
Martin, Thomas E., Juan C. Oteyza, Andy J. Boyce, Penn Lloyd, & Riccardo Ton. (2015). Adult Mortality Probability and Nest Predation Rates Explain Parental Effort in Warming Eggs with Consequences for Embryonic Development Time. The American Naturalist. 186(2). 223–236. 67 indexed citations
7.
Ribeiro, Ângela M., Penn Lloyd, W. R. Dean, Mark Brown, & Rauri C. K. Bowie. (2014). The Ecological and Geographic Context of Morphological and Genetic Divergence in an Understorey-Dwelling Bird. PLoS ONE. 9(2). e85903–e85903. 14 indexed citations
8.
Lloyd, Penn, Fitsum Abadi, Res Altwegg, & Thomas E. Martin. (2014). South temperate birds have higher apparent adult survival than tropical birds in Africa. Journal of Avian Biology. 45(5). 493–500. 35 indexed citations
9.
Ribeiro, Ângela M., Penn Lloyd, Kevin A. Feldheim, & Rauri C. K. Bowie. (2011). Microgeographic socio‐genetic structure of an African cooperative breeding passerine revealed: integrating behavioural and genetic data. Molecular Ecology. 21(3). 662–672. 25 indexed citations
10.
11.
Ribeiro, Ângela M., Penn Lloyd, & Rauri C. K. Bowie. (2011). A TIGHT BALANCE BETWEEN NATURAL SELECTION AND GENE FLOW IN A SOUTHERN AFRICAN ARID-ZONE ENDEMIC BIRD. Evolution. 65(12). 3499–3514. 54 indexed citations
12.
Dijk, René E. van, Ákos Pogány, Jan Komdeur, Penn Lloyd, & Tamás Székely. (2010). Sexual conflict predicts morphology and behavior in two species of penduline tits. BMC Evolutionary Biology. 10(1). 107–107. 11 indexed citations
13.
Ribeiro, Ângela M., Penn Lloyd, Kevin A. Feldheim, & Rauri C. K. Bowie. (2009). Microsatellites in the Karoo scrub‐robin, Cercotrichas coryphaeus (Passeriformes: Muscicapinae): isolation and characterization of 13 autosomal and two sex‐linked loci. Molecular Ecology Resources. 9(2). 636–638. 4 indexed citations
14.
Martin, Thomas E., Sonya K. Auer, Ronald D. Bassar, Alina M. Niklison, & Penn Lloyd. (2007). GEOGRAPHIC VARIATION IN AVIAN INCUBATION PERIODS AND PARENTAL INFLUENCES ON EMBRYONIC TEMPERATURE. Evolution. 61(11). 2558–2569. 209 indexed citations
15.
Lloyd, Penn. (2007). Adult survival, dispersal and mate fidelity in the White‐fronted Plover Charadrius marginatus. Ibis. 150(1). 182–187. 19 indexed citations
16.
Delport, Wayne, Timothy M. Crowe, Penn Lloyd, & Paulette Bloomer. (2006). Population Growth Confounds Phylogeographic Inference in Namaqua Sandgrouse. Journal of Heredity. 98(2). 158–164. 2 indexed citations
17.
Lloyd, Penn, et al.. (2000). The diet and nutrition of the Namaqua sandgrouse, an arid-zone granivore. Journal of Arid Environments. 44(1). 105–122. 5 indexed citations
18.
Lloyd, Penn. (1999). The science and policy behind managing sandgrouse for sustainable utilisation in southern Africa. African Journal of Wildlife Research. 29(2). 35–42. 1 indexed citations
19.
Lloyd, Penn, et al.. (1997). Ecology and genetics of hybrid zones in the southern AfricanPycnonotusbulbul species complex. Ostrich. 68(2-4). 90–96. 12 indexed citations
20.
Hepburn, H. R., R. T. F. Bernard, Bruce Davidson, et al.. (1991). Synthesis and secretion of beeswax in honeybees. Apidologie. 22(1). 21–36. 55 indexed citations

Rankless uses publication and citation data sourced from OpenAlex, an open and comprehensive bibliographic database. While OpenAlex provides broad and valuable coverage of the global research landscape, it—like all bibliographic datasets—has inherent limitations. These include incomplete records, variations in author disambiguation, differences in journal indexing, and delays in data updates. As a result, some metrics and network relationships displayed in Rankless may not fully capture the entirety of a scholar's output or impact.

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